[1
]Faculty of Environmental & Symbiotic Sciences; Prefectural University of Kumamoto;
3-1-100 Tsukide Kumamoto 862-8502 Japan
[2
]Department of Marine Science and Technology; Fukui Prefectural University; 1-1 Gakuen-cho
Obama City Fukui 917-0003 Japan
[3
]National Institute for Environmental Studies; 16-2 Onogawa Tsukuba 305-8506 Japan
[4
]Tohoku National Fisheries Research Institute; 3-27-5 Shinhama-cho Shiogama Miyagi
985-0001 Japan