[1
]Natural Resources and Environmental ScienceUniversity of Nevada Reno1664 N. Virginia
St.RenoNV89557USA
[2
]Program in EcologyEvolution and Conservation BiologyUniversity of Nevada Reno1664
N. Virginia St.RenoNV89557USA
[3
]U.S. Geological SurveyAlaska Science Center4210 University DriveAnchorageAK99508USA