[1
]Laboratory of Plant Systematics and Evolutionary Biology; College of Life Sciences;
Wuhan University; Wuhan; 430072; China
[2
]Botany Department; Jomo Kenyatta University of Agriculture and Technology; Nairobi;
Kenya
[3
]Key Laboratory of Aquatic Botany and Watershed Ecology; Wuhan Botanical Garden; Chinese
Academy of Sciences; Wuhan; 430074; China