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      Additions and corrections to “Family-group names in Coleoptera (Insecta)”



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          Changes to the treatment of Coleoptera family-group names published by Bouchard et al. (2011) are given. These include necessary additions and corrections based on much-appreciated suggestions from our colleagues, as well as our own research. Our ultimate goal is to assemble a complete list of available Coleoptera family-group names published up to the end of 2010 (including information about their spelling, author, year of publication, and type genus). The following 59 available Coleoptera family-group names are based on type genera not included in Bouchard et al. (2011): Prothydrinae Guignot, 1954, Aulonogyrini Ochs, 1953 (Gyrinidae); Pogonostomini Mandl 1954, Merismoderini Wasmann, 1929, †Escheriidae Kolbe, 1880 (Carabidae); Timarchopsinae Wang, Ponomarenko & Zhang, 2010 (Coptoclavidae); Stictocraniini Jakobson, 1914 (Staphylinidae); Cylindrocaulini Zang, 1905, Kaupiolinae Zang, 1905 (Passalidae); Phaeochroinae Kolbe, 1912 (Hybosoridae); Anthypnidae Chalande, 1884 (Glaphyridae); Comophorini Britton, 1957, Comophini Britton, 1978, Chasmidae Streubel, 1846, Mimelidae Theobald, 1882, Rhepsimidae Streubel, 1846, Ometidae Streubel, 1846, Jumnidae Burmeister, 1842, Evambateidae Gistel, 1856 (Scarabaeidae); Protelmidae Jeannel, 1950 (Byrrhoidea); Pseudeucinetini Csiki, 1924 (Limnichidae); Xylotrogidae Schönfeldt, 1887 (Bostrichidae); †Mesernobiinae Engel, 2010, Fabrasiinae Lawrence & Reichardt, 1966 (Ptinidae); Arhinopini Kirejtshuk & Bouchard, 2018 (Nitidulidae); Hypodacninae Dajoz, 1976, Ceuthocera Mannerheim, 1852 (Cerylonidae); Symbiotinae Joy, 1932 (Endomychidae); Cheilomenini Schilder & Schilder, 1928, Veraniini Schilder & Schilder, 1928 (Coccinellidae); Ennearthroninae Chûjô, 1939 (Ciidae); Curtimordini Odnosum, 2010, Mordellochroini Odnosum, 2010 (Mordellidae); Chanopterinae Borchmann, 1915 (Promecheilidae); Heptaphyllini Prudhomme de Borre, 1886, Olocratarii Baudi di Selve, 1875, Opatrinaires Mulsant & Rey, 1853, Telacianae Poey, 1854, Ancylopominae Pascoe, 1871 (Tenebrionidae); Oxycopiini Arnett, 1984 (Oedemeridae); Eutrypteidae Gistel, 1856 (Mycteridae); Pogonocerinae Iablokoff-Khnzorian, 1985 (Pyrochroidae); Amblyderini Desbrochers des Loges, 1899 (Anthicidae); Trotommideini Pic, 1903 (Scraptiidae); Acmaeopsini Della Beffa, 1915, Trigonarthrini Villiers, 1984, Eunidiini Téocchi, Sudre & Jiroux, 2010 (Cerambycidae); Macropleini Lopatin, 1977, Stenopodiides Horn, 1883, Microrhopalides Horn, 1883, Colaphidae Siegel, 1866, Lexiphanini Wilcox, 1954 (Chrysomelidae); †Medmetrioxenoidesini Legalov, 2010, †Megametrioxenoidesini Legalov, 2010 (Nemonychidae); Myrmecinae Tanner, 1966, Tapinotinae Joy, 1932, Acallinae Joy, 1932, Cycloderini Hoffmann, 1950, Sthereini Hatch, 1971 (Curculionidae). The following 21 family-group names, listed as unavailable in Bouchard et al. (2011), are determined to be available: Eohomopterinae Wasmann, 1929 (Carabidae); Prosopocoilini Benesh, 1960, Pseudodorcini Benesh, 1960, Rhyssonotini Benesh, 1960 (Lucanidae); Galbini Beaulieu, 1919 (Eucnemidae); Troglopates Mulsant & Rey, 1867 (Melyridae); Hippodamiini Weise, 1885 (Coccinellidae); Micrositates Mulsant & Rey, 1854, Héliopathaires Mulsant & Rey, 1854 (Tenebrionidae); Hypasclerini Arnett, 1984; Oxaciini Arnett, 1984 (Oedemeridae); Stilpnonotinae Borchmann, 1936 (Mycteridae); Trogocryptinae Lawrence, 1991 (Salpingidae); Grammopterini Della Beffa, 1915, Aedilinae Perrier, 1893, Anaesthetinae Perrier, 1893 (Cerambycidae); Physonotitae Spaeth, 1942, Octotomides Horn, 1883 (Chrysomelidae); Sympiezopinorum Faust, 1886, Sueinae Murayama, 1959, Eccoptopterini Kalshoven, 1959 (Curculionidae). The following names were proposed as new without reference to family-group names based on the same type genus which had been made available at an earlier date: Dineutini Ochs, 1926 (Gyrinidae); Odonteini Shokhin, 2007 (Geotrupidae); Fornaxini Cobos, 1965 (Eucnemidae); Auletobiina Legalov, 2001 (Attelabidae). The priority of several family-group names, listed as valid in Bouchard et al. (2011), is affected by recent bibliographic discoveries or new nomenclatural interpretations. †Necronectinae Ponomarenko, 1977 is treated as permanently invalid and replaced with †Timarchopsinae Wang, Ponomarenko & Zhang, 2010 (Coptoclavidae); Agathidiini Westwood, 1838 is replaced by the older name Anisotomini Horaninow, 1834 (Staphylinidae); Cyrtoscydmini Schaufuss, 1889 is replaced by the older name Stenichnini Fauvel, 1885 (Staphylinidae); Eremazinae Iablokoff-Khnzorian, 1977 is treated as unavailable and replaced with Eremazinae Stebnicka, 1977 (Scarabaeidae); Coryphocerina Burmeister, 1842 is replaced by the older name Rhomborhinina Westwood, 1842 (Scarabaeidae); Eudysantina Bouchard, Lawrence, Davies & Newton, 2005 is replaced by the older name Dysantina Gebien, 1922 which is not permanently invalid (Tenebrionidae). The names Macraulacinae/-ini Fleutiaux, 1923 (Eucnemidae), Anamorphinae Strohecker, 1953 (Endomychidae), Pachycnemina Laporte, 1840 (Scarabaeidae), Thaumastodinae Champion, 1924 (Limnichidae), Eudicronychinae Girard, 1971 (Elateridae), Trogoxylini Lesne, 1921 (Bostrichidae), Laemophloeidae Ganglbauer, 1899 (Laemophloeidae); Ancitini Aurivillius, 1917 (Cerambycidae) and Tropiphorini Marseul, 1863 (Curculionidae) are threatened by the discovery of older names; Reversal of Precedence (ICZN 1999: Art. 23.9) or an application to the International Commission on Zoological Nomenclature will be necessary to retain usage of the younger synonyms. Reversal of Precedence is used herein to qualify the following family-group names as nomina protecta: Murmidiinae Jacquelin du Val, 1858 (Cerylonidae) and Chalepini Weise, 1910 (Chrysomelidae). The following 17 Coleoptera family-group names (some of which are used as valid) are homonyms of other family-group names in zoology, these cases must be referred to the Commission for a ruling to remove the homonymy: Catiniidae Ponomarenko, 1968 (Catiniidae); Homopterinae Wasmann, 1920, Glyptini Horn, 1881 (Carabidae); Tychini Raffray, 1904, Ocypodina Hatch, 1957 (Staphylinidae); Gonatinae Kuwert, 1891 (Passalidae); Aplonychidae Burmeister, 1855 (Scarabaeidae); Microchaetini Paulus, 1973 (Byrrhidae); Epiphanini Muona, 1993 (Eucnemidae); Limoniina Jakobson, 1913 (Elateridae); Ichthyurini Champion, 1915 (Cantharidae); Decamerinae Crowson, 1964 (Trogossitidae); Trichodidae Streubel, 1839 (Cleridae); Monocorynini Miyatake, 1988 (Coccinellidae); Gastrophysina Kippenberg, 2010, Chorinini Weise, 1923 (Chrysomelidae); Meconemini Pierce, 1930 (Anthribidae). The following new substitute names are proposed: Phoroschizus (to replace Schizophorus Ponomarenko, 1968) and Phoroschizidae (to replace Schizophoridae Ponomarenko, 1968); Mesostyloides (to replace Mesostylus Faust, 1894) and Mesostyloidini (to replace Mesostylini Reitter, 1913). The following new genus-group name synonyms are proposed [valid names in square brackets]: Plocastes Gistel, 1856 [Aesalus Fabricius, 1801] (Lucanidae); Evambates Gistel, 1856 [Trichius Fabricius, 1775] (Scarabaeidae); Homoeoplastus Gistel, 1856 [Byturus Latreille, 1797] (Byturidae). Two type genera previously treated as preoccupied and invalid, Heteroscelis Latreille, 1828 and Dysantes Pascoe, 1869 (Tenebrionidae), are determined to be senior homonyms based on bibliographical research. While Dysantes is treated as valid here, Reversal of Precedence (ICZN 1999: Art. 23.9) is used to conserve usage of Anomalipus Guérin-Méneville, 1831 over Heteroscelis.

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