[1
]ISNI 0000 0001 1516 6626, GRID grid.265061.6, Department of Molecular Life Science, Division of Basic Medical Science and Molecular
Medicine, , School of Medicine, Tokai University, ; Isehara, Kanagawa Japan
[2
]ISNI 0000 0001 1516 6626, GRID grid.265061.6, Center for Matrix Biology and Medicine, Graduate School of Medicine, , Tokai University, ; Isehara, Kanagawa Japan
[3
]ISNI 0000 0001 1516 6626, GRID grid.265061.6, The Institute of Medical Sciences, , Tokai University, ; Isehara, Kanagawa Japan
[4
]ISNI 0000 0001 1167 1801, GRID grid.258333.c, Section of Gene Expression Regulation, , Frontier Science Research Center, Kagoshima University, ; Kagoshima, Japan
[5
]ISNI 0000 0004 1762 0759, GRID grid.411951.9, Laboratory Animal Facilities & Services, Preeminent Medical Photonics Education &
Research Center, , Hamamatsu University School of Medicine, ; Hamamatsu, Shizuoka, Japan
[6
]ISNI 0000 0004 0377 284X, GRID grid.415729.c, Division of Molecular Genetics, Shigei Medical Research Institute, ; Minami-ku, Okayama, Japan
[7
]ISNI 0000 0001 1516 6626, GRID grid.265061.6, Department of Applied Biochemistry, School of Engineering, , Tokai University, ; Hiratsuka, Kanagawa Japan
[8
]ISNI 0000 0004 0374 0880, GRID grid.416614.0, Division of Biomedical Engineering, , National Defense Medical College Research Institute, ; Tokorozawa, Saitama, Japan
[9
]GRID grid.410772.7, Department of Bioproduction, , Tokyo University of Agriculture, ; Abashiri, Hokkaido Japan
[10
]ISNI 0000 0001 0666 4105, GRID grid.266813.8, Mouse Genome Engineering Core Facility, Vice Chancellor for Research Office, , University of Nebraska Medical Center, ; Omaha, NE USA
[11
]ISNI 0000 0001 0666 4105, GRID grid.266813.8, Developmental Neuroscience, Munroe Meyer Institute for Genetics and Rehabilitation,
, University of Nebraska Medical Center, ; Omaha, NE USA
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