[1
]Community and Conservation Ecology Group; Centre for Ecological and Evolutionary Studies;
University of Groningen; P.O. Box 11103; 9700 CC Groningen; the Netherlands
[2
]IMARES Texel; P.O. Box 167; 1790 AD; Den Burg; the Netherlands
[3
]National Park Administration of the “Lower Saxon Wadden Sea,”; Virchowstr. 1; 26382;
Wilhelmshaven; Germany