[1
]Behavioural Biology, Department of Biology and Helmholtz Institute, Utrecht University,
Padualaan 8, PO Box 80086, 3508 TB Utrecht, The Netherlands
[2
]Department of Zoology, University of Cambridge, Cambridge CB3 8AA, UK
[3
]School of Biology, University of St Andrews, St Andrews, Fife KY16 9TS, UK