[1
]Department of Life Science; Tunghai University; Taichung Taiwan
[2
]Evolution and Ecology Research Centre; University of New South Wales; Sydney NSW Australia
[3
]Bookend Trust and the School of Biological Sciences; University of Tasmania; Sandy
Bay Tasmania Australia
[4
]Center for Measurement Standards; Industrial Technology Research Institute; Hsinchu
Taiwan
[5
]Center for Tropical Ecology and Biodiversity; Tunghai University; Taichung Taiwan