[1
]Imperial College London; Silwood Park, Buckhurst Road, Ascot Berkshire SL5 7PY United
Kingdom
[2
]Department of Ecology and Evolutionary Biology and Kansas Biological Survey; University
of Kansas; Lawrence Kansas 66047 U.S.A.
[3
]Laboratory of Populations; Rockefeller University; 1230 York Avenue New York New York
10065 U.S.A.