[1
]Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire
naturelle, CNRS, Sorbonne Université , EPHE, CP 50, 57 Rue Cuvier, F–75005, Paris,
France
[2
] Sichuan Cave Exploration Team , 610000, Chengdu, PR China
[3
]Department of Entomology, College of Agriculture, South China Agricultural University
, 510642, Wushan, Guangzhou, PR China