The Tocharians composed verse in hierarchical structures, with the verse dominating major cola, and the major colon in turn dominating one or more minor cola. After providing much-needed descriptive data on Tocharian meter, we assess the evidence for the distinction between major vs. minor caesurae in some of the most popular Tocharian B meters, finding support for the commonly assumed colometries in some but not all cases. Of particular interest is the recurring 4+3-syllable colon, since the violability of its internal (putatively minor) caesura varies significantly across meters. We argue that this varying strictness is indeed a function of the meter as opposed to position in the verse, verse length, idiosyncrasies of certain texts, and so forth. We then use a systematic prose comparison method to test the meters for bridges, finding evidence for monosyllable avoidance in (certain) colon-final positions, despite an overall preference for monosyllables in verse vis-à-vis prose. Finally, we discuss the implications that our study has for the restoration of fragmentary Tocharian texts.