The paradox of the first tier: an agenda for paleobiology

Abstract

Nature's discontinuities occur both in the hierarchical structuring of genealogical individuals and in the distinct processes operating at different scales of time, here called tiers. Conventional evolutionary theory denies this structuring and attempts to render the larger scales as simple extrapolation from (or reduction to) the familiar and immediate—the struggle among organisms at ecological moments (conventional individuals at the first tier). I propose that we consider distinct processes at three separable tiers of time: ecological moments, normal geological time (trends during millions of years), and periodic mass extinctions.