[1
] Department of Zoology and Entomology, Rhodes University, Grahamstown 6140, South
Africa
[2
] Ezemvelo KZN Wildlife, Richards Bay, South Africa
[3
] Department of Botany, Grahamstown, South Africa
[4
] Department of Biology and Mathematics & Statistics, Acadia University, Wolfville,
Nova Scotia, Canada
[5
] Centre for Invasion Biology, South African Institute for Aquatic Biodiversity, South
Africa