[1
]GRID grid.503016.1, ISNI 0000 0001 2160 870X, AMAP, Univ Montpellier, IRD, CNRS, INRAE, CIRAD, ; Montpellier, France
[2
]GRID grid.121334.6, ISNI 0000 0001 2097 0141, CIRAD, UPR Forêts et Sociétés, F-34398 Montpellier, France; Université de Montpellier,
; F-34000 Montpellier, France
[3
]CIRAD, UPR Forêts et Sociétés, Yaoundé, Cameroun
[4
]GRID grid.467700.2, ISNI 0000 0001 2182 2028, Center for Conservation and Sustainability, Smithsonian Conservation Biology Institute,
, National Zoological Park, ; Washington, DC USA
[5
]GRID grid.5342.0, ISNI 0000 0001 2069 7798, CAVELab Computational and Applied Vegetation Ecology, Department of Applied Ecology
and Environmental Biology, Faculty of Bioscience Engineering, , Ghent University, ; Ghent, 9000 Belgium
[6
]FRM Ingénierie, 34130 Mauguio – Grand Montpellier, France
[7
]Institut de pharmacopée et de médecine traditionnelle (Herbier National du Gabon),
CENAREST, Libreville, Gabon
[8
]GRID grid.29273.3d, ISNI 0000 0001 2288 3199, Department of Plant Science, Faculty of Science, , University of Buea, ; Buea, Cameroon
[9
]Terre Environnement Aménagement, 13001 Marseille, France
[10
]GRID grid.4861.b, ISNI 0000 0001 0805 7253, Forest is Life, TERRA Teaching and Research Centre, Gembloux Agro-Bio Tech, , University of Liège, ; Gembloux, Belgium
[11
]GRID grid.412661.6, ISNI 0000 0001 2173 8504, Plant Systematic and Ecology Laboratory (LaBosystE), Department of Biology, Higher
Teachers’ Training College, , University of Yaoundé I, ; P.O. Box 047, Yaoundé, Cameroun
[12
]GRID grid.453560.1, ISNI 0000 0001 2192 7591, ForestGEO, Smithsonian Tropical Research Institute, NMNH - MRC 166, ; P.O. Box 37012, Washington, DC 20013-7012 USA
[13
]Institut de Recherche en Ecologie Tropicale/Centre National de la Recherche Scientifique
et Technologique, Libreville, Gabon
[14
]GRID grid.4989.c, ISNI 0000 0001 2348 0746, Faculty of Sciences, Evolutionary Biology and Ecology, , Université Libre de Bruxelles, CP160/12, 50 Av. F. Roosevelt, ; 1050 Brussels, Belgium
[15
]GRID grid.30064.31, ISNI 0000 0001 2157 6568, School of Biological Sciences, , Washington State University, ; 14204 NE Salmon Creek Ave, Vancouver, WA 98686 USA
License:
Open Access This article is licensed under a Creative Commons Attribution 4.0 International License,
which permits use, sharing, adaptation, distribution and reproduction in any medium
or format, as long as you give appropriate credit to the original author(s) and the
source, provide a link to the Creative Commons license, and indicate if changes were
made. The images or other third party material in this article are included in the
article’s Creative Commons license, unless indicated otherwise in a credit line to
the material. If material is not included in the article’s Creative Commons license
and your intended use is not permitted by statutory regulation or exceeds the permitted
use, you will need to obtain permission directly from the copyright holder. To view
a copy of this license, visit
http://creativecommons.org/licenses/by/4.0/.
The Creative Commons Public Domain Dedication waiver
http://creativecommons.org/publicdomain/zero/1.0/ applies to the metadata files associated with this article.