[1
]GRID grid.4991.5, ISNI 0000 0004 1936 8948, Kennedy Institute of Rheumatology, Nuffield Department of Orthopaedics, Rheumatology
and Musculoskeletal Sciences, , University of Oxford, ; Oxford, UK
[2
]GRID grid.62560.37, ISNI 0000 0004 0378 8294, Center for Data Sciences, Brigham and Women’s Hospital, ; Boston, MA USA
[3
]GRID grid.62560.37, ISNI 0000 0004 0378 8294, Division of Genetics, Department of Medicine, Brigham and Women’s Hospital, ; Boston, MA USA
[4
]GRID grid.38142.3c, ISNI 000000041936754X, Department of Biomedical Informatics, Harvard Medical School, ; Boston, MA USA
[5
]GRID grid.66859.34, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, ; Cambridge, MA USA
[6
]GRID grid.62560.37, ISNI 0000 0004 0378 8294, Division of Rheumatology, Inflammation and Immunity, Department of Medicine, Brigham
and Women’s Hospital and Harvard Medical School, ; Boston, MA USA
[7
]GRID grid.8348.7, ISNI 0000 0001 2306 7492, MRC Weatherall Institute of Molecular Medicine, Radcliffe Department of Medicine,
, University of Oxford, John Radcliffe Hospital, ; Oxford, UK
[8
]GRID grid.8348.7, ISNI 0000 0001 2306 7492, Translational Gastroenterology Unit, NIHR Oxford Biomedical Research Centre, , Oxford University Hospitals NHS Foundation Trust, John Radcliffe Hospital, ; Oxford, UK
[9
]GRID grid.4991.5, ISNI 0000 0004 1936 8948, Jenner Institute, Nuffield Department of Medicine, , University of Oxford, ; Oxford, UK
[10
]GRID grid.4991.5, ISNI 0000 0004 1936 8948, The Wellcome Trust Centre for Human Genetics, , University of Oxford, ; Oxford, UK
[11
]GRID grid.8348.7, ISNI 0000 0001 2306 7492, Department of Paediatrics, John Radcliffe Hospital, ; Oxford, UK
[12
]GRID grid.4991.5, ISNI 0000 0004 1936 8948, Old Road Campus Research Building, Department of Oncology, Medical Sciences Division,
, University of Oxford, ; Oxford, UK
[13
]GRID grid.5379.8, ISNI 0000000121662407, Centre for Genetics and Genomics Versus Arthritis, Manchester Academic Health Science
Centre, , University of Manchester, ; Manchester, UK
[14
]GRID grid.6572.6, ISNI 0000 0004 1936 7486, Rheumatology Research Group, Institute for Inflammation and Ageing, College of Medical
and Dental Sciences, , University of Birmingham, Queen Elizabeth Hospital, ; Birmingham, UK
[15
]GRID grid.412563.7, ISNI 0000 0004 0376 6589, NIHR Birmingham Biomedical Research Centre, , University Hospitals Birmingham NHS Foundation Trust, ; Birmingham, UK
[16
]GRID grid.417570.0, ISNI 0000 0004 0374 1269, Roche Pharma Research and Early Development, Immunology, Infectious Diseases and Ophthalmology
(I2O) Discovery and Translational Area, Roche Innovation Center Basel, ; Basel, Switzerland
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