[1
]Department of Genetics, Evolution and Environment, UCL Genetics Institute, University
College London , London WC1E 6BT, UK
[2
]King’s College London, Social Genetic and Developmental Psychiatry Centre, Institute
of Psychiatry , London SE5 8AF, UK
[3
]Wellcome Trust Centre for Human Genetics, University of Oxford , Oxford OX3 7BN, UK
[4
]Department of Psychology, University of Warwick , Coventry CV4 7AL, UK
[5
]Department of Psychological Sciences, Birkbeck, University of London , London WC1E 7HX, UK
[6
]Department of Psychology, Goldsmiths, University of London , London SE14 6NW, UK
[7
]Department of Psychology and Neuroscience, University of Colorado Boulder , Boulder, Colorado 80309-0345, USA
[8
]Wellcome Trust Sanger Institute , Cambridge CB10 1SA, UK
[9
]Telethon Institute for Child Health Research, Centre for Child Health Research, University
of Western Australia , Crawley, Western Australia, Australia
[10
]Cambridge Institute for Medical Research, University of Cambridge School of Clinical
Medicine , Cambridge CB2 0XY, UK
[11
]UCL Institute of Cognitive Neuroscience, University College London , London WC1N 3AR, UK
[12
]UCL Mental Health Sciences Unit, University College London , London W1W 7EJ, UK
[13
]University of Queensland Diamantia Institute, Translational Research Institute, Princess
Alexandra Hospital, University of Queensland , Brisbane, Queensland QLD 4102, Australia
[14
]Department of Epidemiology and Population Health, London School of Hygiene and Tropical
Medicine , London WC1E 7HT, UK
[15
]Department of Epidemiology and Public Health, University College London , London WC1E 6BT, UK
[16
]Neuropsychiatric Genetics Research Group, Institute of Molecular Medicine, Trinity
College Dublin , Dublin 2, Ireland
[17
]Molecular and Physiological Sciences, The Wellcome Trust , London NW1 2BE, UK
[18
]Centre for Digestive Diseases, Blizard Institute, Queen Mary University of London , London E1 2AT, UK
[19
]Wolfson College , Linton Road, Oxford OX2 6UD, UK
[20
]Peninsula School of Medicine and Dentistry, Associate Deans Office, John Bull Building , Plymouth PL6 8BU, UK
[21
]Clinical Neurosciences, Saint George's University of London , London SW17 0RE, UK
[22
]Department of Medical and Molecular Genetics, King’s College London, King’s Health
Partners Guy’s Hospital , London SE1 9RT, UK
[23
]Biomedical Research Centre, Ninewells Hospital and Medical School , Dundee DD1 9SY, UK
[24
]Department of Clinical Neurosciences, University of Cambridge, Addenbrooke’s Hospital , Cambridge CB2 2QQ, UK
[25
]NIHR Biomedical Research Centre at Moorfields Eye Hospital NHSFT and UCL Institute
of Ophthalmology , London EC1V 2PD, UK
[26
]Department of Molecular Neuroscience, Institute of Neurology, University College London , London WC1N 3BG, UK
[27
]Department of Speech and Hearing Sciences, University of New Mexico , Albuquerque, New Mexico 87131, USA
[28
]Department of Family Science, Institute for Population Research, The Ohio State University , Columbus, Ohio 43210, USA
[29
]Institute for Translational Medicine and Therapeutics, University of Pennsylvania
School of Medicine , Philadelphia, Pennsylvania 19104-5158, USA
[30
]Department of Statistics, University of Oxford , Oxford OX1 3TG, UK
[31
]These authors contributed equally to this work
[32
]These authors jointly supervised this work
License:
This work is licensed under a Creative Commons Attribution 4.0 International License.
The images or other third party material in this article are included in the article’s
Creative Commons license, unless indicated otherwise in the credit line; if the material
is not included under the Creative Commons license, users will need to obtain permission
from the license holder to reproduce the material. To view a copy of this license,
visit http://creativecommons.org/licenses/by/4.0/