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Abstract
Natural selection is conventionally assumed to favour the strong and selfish who maximize
their own resources at the expense of others. But many biological systems, and especially
human societies, are organized around altruistic, cooperative interactions. How can
natural selection promote unselfish behaviour? Various mechanisms have been proposed,
and a rich analysis of indirect reciprocity has recently emerged: I help you and somebody
else helps me. The evolution of cooperation by indirect reciprocity leads to reputation
building, morality judgement and complex social interactions with ever-increasing
cognitive demands.