[1
]Division of Rheumatology, Immunology, and Allergy, Brigham and Women’s Hospital, Harvard
Medical School, Boston, MA, USA.
[2
]Division of Genetics, Brigham and Women’s Hospital, Harvard Medical School, Boston,
MA, USA.
[3
]Program in Medical and Population Genetics, Broad Institute, Cambridge, MA, USA.
[4
]Department of Statistics, Harvard University, Cambridge, MA, USA.
[5
]Centre for Cancer Research, Monash Institute of Medical Research, Monash University,
Clayton, Victoria, Australia.
[6
]Program in Translational NeuroPsychiatric Genomics, Institute for the Neurosciences,
Department of Neurology, Brigham and Women’s Hospital, Boston, MA, USA.
[7
]Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto,
Japan.
[8
]Department of Rheumatology and Clinical immunology, Graduate School of Medicine, Kyoto
University, Kyoto, Japan.
[9
]Institute of Rheumatology, Tokyo Women’s Medical University, Tokyo, Japan.
[10
]Laboratory for Autoimmune Diseases, Center for Integrative Medical Sciences, RIKEN,
Yokohama, Japan.
[11
]Immunology Biomarkers Group, Genentech, South San Francisco, CA, USA.
[12
]Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville,
TN, USA.
[13
]Department of Medicine, Vanderbilt University School of Medicine, Nashville, TN, USA.
[14
]New York University Hospital for Joint Diseases, New York, NY, USA.
[15
]Department of Medicine, Albany Medical Center and The Center for Rheumatology, Albany,
NY, USA.
[16
]Division of Rheumatology, Department of Medicine, New York, Presbyterian Hospital,
College of Physicians and Surgeons, Columbia University, New York, NY, USA.
[17
]Department of Rheumatology and Immunology, Shanghai Changzheng Hospital, Second Military
Medical University, Shanghai 200003, China.
[18
]Department of Pharmacology, Second Military Medical University, Shanghai, 200433,
China
[19
]University of Queensland Diamantina Institute, Translational Research Institute, Brisbane,
Queensland, Australia.
[20
]Queensland Brain Institute, The University of Queensland, Brisbane, Queensland, Australia.
[21
]Samuel Lunenfeld Research Institute, Mount Sinai Hospital, Toronto, Canada.
[22
]Toronto General Research Institute, Toronto, Canada.
[23
]Department of Medicine, University of Toronto, Toronto, Canada.
[24
]Department of Medicine, Mount Sinai Hospital and University of Toronto, Toronto, Canada.
[25
]Department of Genetics, University Medical Center Groningen, University of Groningen,
Hanzeplein 1, Groningen, the Netherlands.
[26
]Estonian Genome Center, University of Tartu, Riia 23b, 51010, Tartu, Estonia.
[27
]Division of Endocrinology, Children’s Hospital, Boston, MA, USA.
[28
]School of Computer and Information Technology, Beijing Jiaotong University, Beijing,
China.
[29
]The Department of Psychiatry at Mount Sinai School of Medicine, New York, USA.
[30
]Department of Human Genetics, Radboud University Nijmegen Medical Centre, Nijmegen,
The Netherlands.
[31
]Department of Rheumatology, Radboud University Nijmegen Medical Centre, Nijmegen,
The Netherlands.
[32
]Department of Rheumatology and Clinical Immunology, Arthritis Center Twente, University
Twente & Medisch Spectrum Twente, Enschede, The Netherlands.
[33
]Department of Clinical Pharmacy and Toxicology, Leiden University Medical Center,
Leiden, The Netherlands.
[34
]Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands.
[35
]Service de Rhumatologie et INSERM U699 Hôpital Bichat Claude Bernard, Assistance Publique
des Hôpitaux de Paris, Paris, France.
[36
]Université Paris 7-Diderot, Paris, France.
[37
]Université Paris-Sud, Orsay, France.
[38
]APHP–Hôpital Bicêtre, INSERM U1012, Le Kremlin Bicêtre, Paris, France.
[39
]Division of Clinical Immunology and Rheumatology, Department of Medicine, University
of Alabama at Birmingham, Birmingham, AL, USA.
[40
]AMC/University of Amsterdam, Amsterdam, the Netherlands.
[41
]GlaxoSmithKline, Stevenage, U.K.
[42
]University of Cambridge, Cambridge, U.K.
[43
]Institut National de la Santé et de la Recherche Médicale (INSERM) U1012.
[44
]Université Paris-Sud 11, Rhumatologie, Hôpital Bicêtre, Assistance Publique-Hôpitaux
de Paris (AP-HP), Le Kremlin Bicêtre, France.
[45
]Department of Rheumatology, Hanyang University Hospital for Rheumatic Diseases, Seoul,
South Korea.
[46
]Instituto de Parasitologia y Biomedicina Lopez-Neyra, CSIC, Granada, Spain.
[47
]Department of Rheumatology, Hospital Marques de Valdecilla, IFIMAV, Santander, Spain.
[48
]Hospital Clinico San Carlos, Madrid, Spain.
[49
]Department of Public Health and Clinical Medicine, Umeå University, Umeå, Sweden.
[50
]Department of Rheumatology, Umeå University, Umeå, Sweden.
[51
]Channing Laboratory, Department of Medicine, Brigham and Women’s Hospital, Harvard
Medical School, Boston, MA, USA.
[52
]Section of Rheumatology, Boston University School of Medicine, Boston, MA, USA.
[53
]Clinical Epidemiology Research and Training Unit, Boston University School of Medicine,
Boston, MA, USA.
[54
]Centre d’Etude du Polymorphisme Humain (CEPH), Paris, France.
[55
]Université Paris 13 Sorbonne Paris Cité, UREN (Nutritional Epidemiology Research Unit),
Inserm (U557), Inra (U1125), Cnam, Bobigny, France.
[56
]McGill University and Génome Québec Innovation Centre, Montréal, Canada.
[57
]Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University
of Manchester, Manchester Academic Health Science Centre, Manchester, UK.
[58
]National Institute for Health Research, Manchester Musculoskeletal Biomedical Research
Unit, Central Manchester University Hospitals National Health Service Foundation Trust,
Manchester Academic Health Sciences Centre, Manchester, UK.
[59
]Department of Clinical Immunology and Rheumatology & Department of Genome Analysis,
Academic Medical Center/University of Amsterdam, Amsterdam, The Netherlands.
[60
]Division of Rheumatology and Clinical Immunology, University of Pittsburgh, Pittsburgh,
PA, USA.
[61
]Rosalind Russell Medical Research Center for Arthritis, Division of Rheumatology,
Department of Medicine, University of California San Francisco, San Francisco, CA,
USA.
[62
]Unit of Statistical Genetics, Center for Genomic Medicine Graduate School of Medicine
Kyoto University, Kyoto, Japan.
[63
]Laboratory for Genotyping Development, Center for Integrative Medical Sciences, RIKEN,
Yokohama, Japan.
[64
]Rheumatology Unit, Department of Medicine (Solna), Karolinska Institutet, Stockholm,
Sweden.
[65
]The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health
System, Manhasset, NY, USA.
[66
]NIHR Manchester Musculoskeletal Biomedical, Research Unit, Central Manchester NHS
Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK.
[67
]Section of Genetic Medicine, University of Chicago, Chicago, IL, USA.
[68
]Institute for Genomics and Systems Biology, University of Chicago, Chicago, IL, USA.
[69
]Laboratory for Statistical Analysis, Center for Integrative Medical Sciences, RIKEN,
Yokohama, Japan.
[70
]Core Research for Evolutional Science and Technology (CREST) program, Japan Science
and Technology Agency, Kawaguchi, Saitama, Japan.
[71
]Institut National de la Sante et de la Recherche Medicale (INSERM) Unite U852, Kyoto
University Graduate School of Medicine, Kyoto, Japan.
[72
]Department of Allergy and Rheumatology, Graduate School of Medicine, the University
of Tokyo, Tokyo, Japan.
