[1
] National Institute of Agrobiological Resources, Tsukuba, Ibaraki 305-8602, Japan
[2
] Bio-oriented Technology Research Advancement Institution, Omiya, Saitama 331-8537,
Japan
[3
] Institute of the Society for Techno-innovation of Agriculture, Forestry and Fisheries,
Tsukuba, Ibaraki 305-0854, Japan
[4
] Department of Biology, Faculty of Science, Toyama University, Toyama, 930-8555, Japan