[1
]Department of Biology, University of Joensuu, PO Box 111, FIN-80101 Joensuu, Finland
[2
]Department of Biology, University of Joensuu, PO Box 111, FIN-80101 Joensuu, Finland;
Department of Geography and the Centre for Climate and Global Change Research, McGill
University, 805 Sherbrooke Street West, Montreal, Quebec, H3A 2K6, Canada