[1
]Department of Wildlife, Fish, and Environmental Studies Swedish University of Agricultural
Sciences Umeå SE‐90183 Sweden
[2
]Norwegian Institute for Nature Research Sognsveien 68 Oslo NO‐0855 Norway
[3
]Department of Zoology Centre for African Conservation Ecology Nelson Mandela University
Port Elizabeth 6031 South Africa
[4
]Copernicus Institute of Sustainable Development Environmental Sciences Utrecht University
Utrecht 3548 The Netherlands
[5
]Norwegian Institute for Nature Research PO Box 5685 Torgard Trondheim NO‐7485 Norway
[6
]Department of Ecology Swedish University of Agricultural Sciences Grimsö Wildlife
Research Station RiddarhyttanSE‐73993 Sweden
[7
]Department of Forestry and Wildlife Management Inland Norway University of Applied
Sciences Koppang NO‐2480 Norway