[1
]Ministry of Education Key Laboratory for Biodiversity and Ecological Engineering;
College of Life Sciences; Beijing Normal University; Beijing China
[2
]Key Laboratory of Zoological Systematics and Evolution; Institute of Zoology; Chinese
Academy of Sciences; Chaoyang District Beijing 100101 China
[3
]Swedish Species Information Centre; Swedish University of Agricultural Sciences; Box
7007 SE-750 07 Uppsala Sweden
[4
]Section of Systematics and Biodiversity, Biology and Environmental Sciences; University
of Gothenburg; Box 463 SE-405 30 Gothenburg Sweden
[5
]Yamashina Institute for Ornithology; 115 Konoyama Abiko Chiba 270-1145 Japan