[1
]Laboratory for Marine Fisheries and Aquaculture; Qingdao National Laboratory for Marine
Science and Technology; Key Laboratory of Maricultural Organism Disease Control; Ministry
of Agriculture; Yellow Sea Fisheries Research Institute; Chinese Academy of Fishery
Sciences; Qingdao China
[2
]College of Fishery and Life Sciences; Shanghai Ocean University; Shanghai China
[3
]Marine Science and Engineering College; Qingdao Agricultural University; Qingdao China